The novel The Saint Beloved by the Prince is set in the land of Lacia, where spirits control the elements, each of which is overseen by a "Spirit King." On rare occasions, these Spirit Kings may bless a single human, granting them immense magical control over their respective element. The heroine, Amelia Logan, and the villainess, Claudia Leitzya, are two sides of the same coin, blessed by the Light and Dark Spirit Kings respectively. However, while Amelia is beloved, Claudia is hated and rejected due to her dark skin, ebony locks, and aptitude for dark magic.

After dying in modern-day Japan, Sakura is reborn as an infant Claudia. While difficult at first, Claudia tries to adjust to this new life, all the while accompanied by the Dark Spirit King Gerald. But even this task is far from easy; people fear her at first glance, no matter how she acts toward them. Still, Claudia is set on changing her fate from the novel, proving she is a good person and breaking free from the shackles of a "villainess."